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会计学Microbescanbedividedintometabolicclasseswhichrelatetothesourcesofenergytheyuse.Thethreegroupsare: 1.heterotrophswhichutilizeorganicmoleculesasasourceofenergy(thesearealsocalledchemo-organotrophs化能有机营养型); 2.phototrophswhichobtainenergyfromlight 3.lithotrophswhichobtainenergyfrominorganiccompounds. Carbon(碳素)forcellsynthesisisobtainedfromorganicmolecules;however,somemicrobes,includingthephototrophs,fixCO2.Catabolism Metabolism anabolism ① complexmoleculesimplemolecules+ATP+[H]* (Organicmaterial)② *[H]standsforreducingpower(还原力) ①catabolismenzymecomplex(分解代谢酶系)②anabolismenzymecomplex(合成代谢酶系)organicmaterial Primaryenergysources(最初能源)rayirradiation(light) Chemoheterotrophsreducedinorganicmaterial Phototrophs chemoautotrophs universalenergysource(通用能源)——ATPSectionA Heterotrophicpathways Biologicaloxidationisreferredtoasalltheenergy-producingoxidativereactionsinthelivingorganism.Definition:Aseriesofenergy-producingoxidativereactionsoccursinthelivingcellsarecalledbiologicaloxidation.Functionofbiologicaloxidation(生物氧化的功能) Produceenergy(产能) Producereducingpower[H](产还原力) Smallintermediatemetabolites(产小分子中间代谢物)Pathwaysofremovingofhydrogen:(inthecaseofGlucose) EMP(glycosis) HMP(pentosephosphatepathwayorWD) ED(KDPG) TCAcycle(citricacidcycle)removeMostmicrobesutilizetheglycolytic(糖酵解)pathwayforthecatabolismofcarbohydratessuchasglucoseandfructose.Theproductsofthispathwayarepyruvate,whichcanbefurthermetabolizedviathecitricacidcycle,formingadenosine5’-triphosphate(ATP)andthereducedformofnicotinamideadeninedinucleotide(烟酰胺腺嘌呤核苷酸NADH+H+.)Thispathwayislocatedinthecytoplasmofmicrobesandcanfunctioninthepresenceorabsenceofoxygen.ThebacterialgeneraPseudomonas,Rhizobium(根瘤菌)andAgrobacter(农杆菌)substitutetheEntner-Doudoroffpathwayfortheglycolyticpathway.Thispathwayisnotasefficientinproducingenergy,with1moleofATPbeingformedforeachmoleofglucosemetabolized.ThepentosephosphatepathwayorhexosemonophosphatepathwaymayoperateatthesametimeasglycolysisortheEntner-Doudoroffpathway.Thispathwaycanalsoo